Cyprus Turtles

Research interests have always been extremely broad, as is the nature of marine turtle research as even now so much is yet to be understood. A legacy of peer-reviewed publications has been built by Marine Turtle Research Group (MTRG) from studies conducted in North Cyprus.

Below you may download the publications written by the data gathered or studies conducted in North Cyprus. To download please click on the icon next to the publications textual information.

Duncan EM, Akbora HD, Baldi P, Beton D, Broderick AC, Çicek BA, Crowe-Harland C, Davey C, DeSerisy T, Fuller WJ, Haywood JC, Hsieh YJ, Kaya E, Omeyer LCM, Ozkan M, Palmer JL, Roast E, Santillo D, Schneider J, Snape RTE, Sutherland KC, Godley BJ (2024) Marine turtles as bio- indicators of plastic pollution in the eastern Mediterranean. Mar. Pollut. Bull. 201, 116141

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Snape RTE, Beton D, Broderick AC, Omeyer LC, Godley BJ (2024) Flashing NetLights reduce bycatch in small-scale fisheries of the Eastern Mediterranean. Fish. Res. 272, 106919

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Fuentes M, Santos, A, Abreu‐Grobois A, Briseño‐Dueñas R, Al‐Khayat J, Hamza S, Saliba S, Anderson D, Rusenko KW, Mitchell NJ, Gammon M, Bentley BP, Beton D, Booth DTB, Broderick AC, Colman L, Snape RTE, … JR Monsinjon (2024) Adaptation of sea turtles to climate warming: Will phenological responses be sufficient to counteract changes in reproductive output? Glob. Change Biol. 30, e16991

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Palmer JL, Armstrong C, Akbora HD, Beton D, Çağlar Ç, Godley BJ, Metcalfe K, Özkan M, Snape RTE, Broderick AC (2024) Behavioural patterns, spatial utilization and landings composition of a small-scale fishery in the eastern Mediterranean. Fish. Res. 269, 106861

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Kaska A, Koc G, Sozbilen D, Salih D, Glidan A, Elsowayeb AA, Saied A, Margaritoulis D, Theodorou P, Rees A, Snape R, Broderick A, Godley B, Beton D, Ozkan M, Jribi I, Ismail MB, Hmida AB, Badreddine A, Sacdanaku E, Rae V, Abdelwarith MS, Naguib N, Nakhla LB, Limam A, Abderrahim M, Sémelin J, Kaska Y (2023) Increased sample size provides novel insights into population structure of Mediterranean loggerhead sea turtles. Conservation Genetics Res. 15(1-2), pp. 67-75

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O’Keefe M, Bengil EGT, Palmer JL, Beton D, Çağlar Ç, Godley BJ, Özkan M, Snape RTE, Broderick AC (2023) Diversity and distribution of elasmobranchs in the coastal waters of Cyprus: using bycatch data to inform management and conservation. Front. Mar. Sci. 1181437

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de Kock W, Mackie M, Ramsøe M, Allentoft ME, Broderick AC, Haywood JC, Godley BJ, Snape RTE, Bradshaw PJ, Genz H, von Tersch M, Dee MW, Palsbøll PJ, Alexander M, Taurozzi AJ, Çakırlar C (2023) Threatened North African seagrass meadows have supported green turtle populations for millennia. PNAS. 120, e2220747120

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Çelik S, Beton D, Çiçek BA, Snape RTE, Başkale E (2023) Metal accumulation in juvenile and sub-adult loggerhead and green turtles in Northern Cyprus. Environ. Pollut. 316, 120482

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Kot CY, Akesson S, Alfaro-Shigueto J, Llanos DF, Antonopoulou M, Balazs GH, Baverstock WR, Blumenthal JM, Broderick AC, Bruno I, Canbolat AF, Casale P, Cejudo D, Coyne MS, Curtice C, DeLand S, DiMatteo A, Dodge K, Dunn DC, Esteban N, Formia A, Fuentes MMPB, Fujioka E, Garnier J, Godfrey MH, Godley BJ, Carman VG, Harrison A, Hart CE, Hawkes LA, Hays GC, Hill N, Hochscheid S, Kaska Y, Levy Y, Ley-Quinonez CP, Lockhart GG, Lopez-Mendilaharsu M, Luschi P, Mangel JC, Margaritoulis D, Maxwell SM, McClellan CM, Metcalfe K, Mingozzi A, Moncada FG, Nichols WJ, Parker DM, Patel SH, Pilcher NJ, Poulin S, Read AJ, Rees AF, Robinson DP, Robinson NJ, Sandoval-Lugo AG, Schofield G, Seminoff JA, Seney EE, Snape RTE, Sozbilen D, Tomas J, Varo-Cruz N, Wallace BP, Wildermann NE, Witt MJ, Zavala-Norzagaray AA, Halpin PN (2022) Network analysis of sea turtle movements and connectivity: A tool for conservation prioritization. Divers. Distrib. 28, 810-829

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Snape RTE, Beton D, Davey S, Godley BJ, Haywood J, Omeyer LC, Özkan M, Broderick AC (2022) Mediterranean green turtle population recovery increasingly depends on Lake Bardawil, Egypt. GECCO 40, e02336

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Darmon G, Schulz M, Matiddi M, Loza AL, Tomas J, Camedda A, Chaieb O, Hili HAE, Bradai MN, Bray L, Claro F, Dellinger T, Dell’Amico F, Lucia GA, Duncan EM, Gambaiani D, Godley BJ, Kaberi H, Kaska Y, Martin J, Moreira C, Ostiategui P, Pham CK, Piermarini R, Revuelta O, Rodriguez Y, Silvestri C, Snape RTE, Sozbilen D, Tsangaris C, Vale M, Vandeperre F, Miaud C (2022) Drivers of litter ingestion by sea turtles: Three decades of empirical data collected in Atlantic Europe and the Mediterranean. Mar

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DiMatteo A, Cañadas A, Roberts J, Sparks L, Panigada S, Boisseau O, Moscrop A, Fortuna CM, Lauriano G, Holcer D, Peltier H, Ridoux V, Raga JA, Tomas J, Broderick AC, Godley BJ, Haywood J, March D, Snape R, Sagarminaga R, Hochscheid S (2022) Basin- wide estimates of loggerhead turtle abundance in the Mediterranean Sea derived from line transect surveys. Front. Mar. Sci. 9:930412

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Papageorgiou M, Bengil EGT, Snape R, Hadjioannou (2022) Increased knowledge affects public attitude and perception towards elasmobranchs and support for conservation. MediterrMarSci 23:637–649

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Snape RTE, Akbora HD, Çiçek BA, Kaya E, Özkan M, Palmer J, Beton D (2022) Spate of loggerhead turtle strandings suggest predation by Mediterranean monk seal. MedTurtleBul 1: 37-43

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Beton D, Broderick AC, Godley BJ, Kolac E, Ok M, Snape RTE (2021) New monitoring confirms regular breeding of the Mediterranean monk seal in Northern Cyprus. Oryx 55:522-525

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Omeyer LCM, Stokes KL, Beton D, Çiçek BA, Davey S, Fuller WJ, Godley BJ, Sherley RB, Snape RTE, Broderick, AC (2021) Investigating differences in population recovery rates of two sympatrically nesting sea turtle species. AnimConserv 24:832-846

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Palmer JL, Beton D, Cicek BA, Davey S, Duncan EM Fuller WJ, Godley BJ, Haywood JC, Huseyinoglu MF, Omeyer LCM, Scheider MJ, Snape RTE, Broderick AC (2021) Dietary analysis of two sympatric marine turtle species in the eastern Mediterranean. MarBiol 168:94

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Edwards RC, Godley BJ, Nuno A (2020) Exploring connections among the multiple outputs and outcomes emerging from 25 years of sea turtle conservation in Northern Cyprus. JNatureConserv 55:125816

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Haywood JC, Casale P, Freggi D, Fuller WJ, Godley BJ, Lazar B, Margaritoulis D, Rees AF, Shutler JD, Snape RT, Swain-Diaz NR, Widdicombe S, Broderick AC (2020) Foraging ecology of Mediterranean juvenile loggerhead turtles: insights from C and N stable isotope ratios. MarBiol 167:28

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Haywood JC, Fuller WJ, Godley BJ, Margaritoulis D, Shutler JD, Snape RTE, Widdicombe S, Zbinden J, Broderick AC (2020) Spatial ecology of loggerhead turtles: insights from stable isotope markers and satellite telemetry. DiversityDist 26:368-381

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Duncan EM, Arrowsmith JAA, Bain CE, Bowdery H, Broderick AC, Chalmers T, Fuller WJ, Galloway TS, Lee JH, Lindeque PK, Omeyer LCM, Snape RTE, Godley BJ (2019). Diet-related selectivity of macroplastic ingestion in green turtles (Chelonia mydas) in the eastern Mediterranean. Sci Rep 9

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Haywood JC, Fuller WJ, Godley BJ, Shutler JD, Widdicombe S, Broderick AC (2019). Global review and inventory: how stable isotopes are helping us understand ecology and inform conservation of marine turtles. MarEcolProgSer 613: 217–245

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Omeyer LCM, Casale P, Fuller WJ, Godley BJ, Holmes KE, Snape RTE, Broderick AC (2019). The importance of passive integrated transponder (PIT) tags for measuring life-history traits of sea turtles. Elsevier 240

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Omeyer LCM, Fuller WJ, Godley BJ, Snape RTE, Broderick AC (2019). The effect of biologging systems on reproduction, growth and survival of adult sea turtles. Mov. Ecol. 7

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Bradshaw PJ, Broderick AC, Carreras C, Fuller W, Snape RTE, Wright LI, Godley BJ (2018). Deﬁning conservation units with enhanced molecular tools to reveal ﬁne scale structuring among Mediterranean green turtle rookeries. BiolCons 222:253-260

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Casale P, Broderick A, Camiñas JA, Cardona L, Carreras C, Demetropoulos A, Fuller WJ, Godley BJ, Hochscheid, Kaska Y, Lazar B, Margaritoulis D, Panagopoulou A, Rees AF, Tomás J, Türkozan O (2018). Mediterranean sea turtles: current knowledge and priorities for conservation and research. EndangSpeciesRes 36: 229–267

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Duncan EM, Arrowsmith J, Baina C, Broderick AC, Lee J, Metcalfe K, Pikesley S, Snape RTE, van Sebilled E, Godley BJ (2018). The true depth of the Mediterranean plastic problem: Extreme microplastic pollution on marine turtle nesting beaches in Cyprus. MarinePollBull 136: 334-340

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Duncan EM, Broderick AC, Fuller WJ, Galloway TS, Godfrey MH, Hamann M, Limpus C, Lindeque PK, Mayes AG, Omeyer LCM, Santillo D, Snape RTE, Godley BJ (2018). Microplastic ingestion ubiquitous in marine turtles. GlobalChangeBiol 25: 744-752

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Omeyer, LCM, Fuller WJ, Godley BJ, Snape RTE, Broderick AC (2018). Determinate or indeterminate growth? Revisiting the growth strategy of sea turtles. MarEcolProgSer 596: 199–211

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Snape RTE, Bradshaw PJ, Broderick AC, Fuller WJ, Stokes KL, Godley BJ (2018). Oﬀ-the-shelf GPS technology to inform marine protected areas for marine turtles. BiolCons 227: 301-309

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Snape RTE, Broderick AC, Çiçek BA, Fuller WJ, Tregenza N, Witt M, Godley BJ (2018). Conflict between Dolphins and a data-scarce fishery of the European Union. HumEcol 46:423–433

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Tikochinski Y, Bradshaw PJ, Mastrogiacomo A, Broderick AC, Daya A, Demetropoulos A, Demetropoulos S, Eliades N, Fuller WJ, Godley BJ, Kaska Y, Levy L, Snape RTE, Wright LI, Carrera C (2018). Mitochondrial DNA short tandem repeats unveil hidden population structuring and migration routes of an endangered marine turtle. AquatConserv 28: 788-797

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Varela MR, Patrício AR, Anderson K, Broderick AC, DeBell L, Hawkes LA, Tilley D, Snape RTE, Westoby MJ, Godley BJ (2018). Assessing climate change associated sea‐level rise impacts on sea turtle nesting beaches using drones, photogrammetry and a novel GPS system. GlobalChangeBiol 25: 753-762

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Bradshaw PJ, Broderick AC, Carreras C, Inger R, Fuller W, Snape RTE, Stokes KL, Godley BJ (2017). Satellite tracking and stable isotope analysis highlight differential recruitment among foraging areas in green turtles. MarEcolProgSer 582:201-214

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Duncan EM, Botterell ZLR, Broderick AC, Galloway TS, Lindeque PK, Nuno A, Godley BJ (2017). A global review of marine turtle entanglement in anthropogenic debris: a baseline for further action. EndangSpecRes 34:431-448

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Omeyer LCM, Godley BJ, Broderick AC (2017). Growth rates of adult sea turtles. EndangSpecRes 34:357-371

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Snape RTE, Broderick AC, Cicek BA, Fuller WJ, Glen F, Stokes K, Godley BJ (2016). Shelf life: neritic habitat use of a turtle population highly threatened by fisheries. DiversityDistrib :1-11

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Snape RTE (2015). Artisanal fishers help to map marine turtle distribution and bycatch: further evidence for an important foraging ground in Famagusta Bay, North Cyprus Testudo 8:51-59

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Stokes KL, Broderick AC, Canbolat AF, Candan O,Fuller WJ, Glen F, Levy Y, Rees AF, Rilov G, Snape RT, Stott I, Tchernov D, Godley BJ (2015). Migratory corridors and foraging hotspots: critical habitats identified for Mediterranean green turtles DiversityDistrib 21:665-674

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Casale P, Freggi D, Furii G, Vallini C, Salvemini P, Deflorio M, Totaro G, Raimondi S, Fortuna C, Godley BJ (2014). Annual survival probabilities of juvenile loggerhead sea turtles indicate high anthropogenic impact on Mediterranean populations AquatConserv:

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Horne CR, Fuller WJ, Godley BJ, Rhodes KA, Snape R, Stokes KL, Broderick AC (2014). The effect of thermal variance on the phenotype of marine turtle offspring PhysiolBiochemZool 87:796-804

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Stokes KL, Fuller WJ, Glen F, Godley BJ, Hodgson DJ, Rhodes KA, Snape RTE, Broderick AC (2014). Detecting green shoots of recovery: the importance of long-term individual-based monitoring of marine turtles AnimConserv 17:593-602

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Fuller WJ, Godley BJ, Hodgson DJ, Reece SE, Witt MJ, Broderick AC (2013). Importance of spatio-temporal data for predicting the effects of climate change on marine turtle sex ratios MarEcolProgSer 488:267-274

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Snape RTE, Beton D, Broderick AC, Cicek BA, Fuller WJ, Ozden O, Godley BJ (2013). Strand monitoring and anthropological surveys provide insight into marine turtle bycatch in small-scale fisheries of the eastern Mediterranean ChelonConservBiol 12:44-55

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Wright LI, Fuller WJ, Godley BJ, McGowan A, Tregenza T, Broderick AC (2013). No benefits of polyandry to female green turtles BehaEco 24:1022-1029

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Casale P, Broderick AC, Freggi D, Mencacci R, Fuller WJ, Godley BJ, Luschi P (2012). Long-term residence of juvenile loggerhead turtles to foraging grounds: a potential conservation hotspot in the Mediterranean AquatConserv 22:144-154

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Wright LI, Fuller WJ, Godley BJ, McGowan A, Tregenza T, Broderick AC (2012). Reconstruction of paternal genotypes over multiple breeding seasons reveals male green turtles do not breed annually MolEcol 21:3625-3635

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Scott R, Hodgson DJ, Witt MJ, Coyne MS, Adnyana W, Blumenthal JM, Broderick AC, Canbolat AF, Catry P, Ciccione S, Delcroix E, Hitipeuw C, Luschi P, Pet-Soede L, Pendoley K, Richardson PB, Rees AF, Godley BJ (2012). Global analysis of satellite tracking data shows that adult green turtles are significantly aggregated in Marine Protected Areas GlobalEcolBiogeogr 21:1053–1061

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Wright LI, Stokes KL, Fuller WJ, Godley BJ, McGowan A, Snape R, Tregenza T, Broderick AC (2012). Turtle mating patterns buffer against disruptive effects of climate change ProcRSocB 279:2122-7

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Fuller WJ, Broderick AC, Enever R, Thorne P, Godley BJ (2010). Motile homes: a comparison of the spatial distribution of epibiont communities on Mediterranean sea turtles JNatHist 44:1743-1753

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Hamann M, Godfrey MH, Seminoff JA, Arthur K, Barata PCR, Bjorndal KA, Bolten AB, Broderick AC, Campbell LM, Carreras C, Casale P, Chaloupka M, Chan SKF, Coyne MS, Crowder LB, Diez CE, Dutton PH, Epperly SP, FitzSimmons NN, Formia A, Girondot M, Hays (2010). Global research priorities for sea turtles: informing management and conservation in the 21st century EndangSpecRes 11:245-269

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Witt MJ, Hawkes LA, Godfrey MH, Godley BJ, Broderick AC (2010). Predicting the impacts of climate change on a globally distributed species: the case of the loggerhead turtle JExpBiol 213:901-911

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Fuller WJ, Broderick AC, Hooker SK, Witt MJ, Godley BJ (2009). Insights into habitat utilization by green turtles (Chelonia mydas) during the inter-nesting period using animal-borne digital cameras MarTechnolSocJ 43:1-9

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Fuller WJ, Broderick AC, Phillips RA, Silk JRD, Godley BJ (2008). Utility of geolocating light loggers for indicating at-sea movements in sea turtles EndangSpecRes 4:139-146

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Jackson AL, Broderick AC, Fuller WJ, Glen F, Ruxton GD, Godley BJ (2008). Sampling design and its effect on population monitoring: How much monitoring do turtles really need? BiolConserv 141:2932-2941

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Broderick AC, Coyne MS, Fuller WJ, Glen F, Godley BJ (2007). Fidelity and over-wintering of sea turtles ProcRSocB 274:1533-1538

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Godley BJ, Blumenthal JM, Broderick AC, Coyne MS, Godfrey MH, Hawkes LA, Witt MJ (2007). Satellite tracking of sea turtles: where have we been and where do we go next EndangSpecRes 3:1-20

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Blumenthal JM, Solomon JL, Bell CD, Austin TJ, Ebanks-Petrie SG, Coyne MS, Broderick AC, Godley BJ (2006). Satellite tracking highlights the need for international cooperation in marine turtle management EndangSpecRes 7:1-11

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Broderick AC, Frauenstein R, Glen F, Hays GC, Jackson AL, Pelembe T, Ruxton GD, Godley BJ (2006). Are green turtles globally endangered? Global Ecology and Biogeography 15:21-26

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Coyne MS, Godley BJ (2005). Satellite Tracking and Analysis Tool (STAT): an integrated system for archiving, analyzing and mapping animal tracking data MarEcolProgSer 127:1-7

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Glen F, Broderick AC, Godley BJ, Hays GC (2005). Patterns in the emergence of green (Chelonia mydas) and loggerhead (Caretta caretta) turtle hatchlings from their nests MarBiol 146:1039-1049

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Broderick AC, Glen F, Godley BJ, Hays GC (2003). Variation in reproductive output of marine turtles JExpMarBiolEcol 288:95-109

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Glen F, Broderick AC, Godley BJ, Hays GC (2003). Incubation environment affects phenotype of naturally incubated green turtle hatchlings JMarBiolAssUK 83:1183-1186

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Godley BJ, Broderick AC, Glen F, Hays GC (2003). Post-nesting movements and submergence patterns of loggerhead marine turtles in the Mediterranean assessed by satellite tracking JExpMarBiolEcol 287:119-134

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Reece SE, Broderick AC, Godley BJ, (2002) SA West Evolutionary Ecology Research 4 (5), 737-748

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Broderick AC, Glen F, Godley BJ, Hays GC (2002). Estimating the number of green and loggerhead turtles nesting annually in the Mediterranean Oryx 36:227-235

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Godley BJ, Richardson S, Broderick AC, Coyne MS, Glen F, Hays GC (2002). Long-term satellite telemetry of the movements and habitat utilisation by green turtles in the Mediterranean Ecography 25:352-362

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Hays GC, Broderick AC, Glen F, Godley BJ, Houghton JDR, Metcalfe JD (2002). Water temperature and internesting intervals for loggerhead (Caretta caretta) and green (Chelonia mydas) sea turtles JThermBiol 27:429-432

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Hays GC, Glen F, Broderick AC, Godley BJ, Metcalfe JD (2002). Behavioural plasticity in a large marine herbivore: contrasting patterns of depth utilisation between two green turtle (Chelonia mydas) populations MarBiol 141:985-990

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Houghton JDR, Broderick AC, Godley BJ, Metcalfe JD, Hays GC (2002). Diving behaviour during the internesting interval for loggerhead turtles Caretta caretta nesting in Cyprus MarEcolProgSer 227:63-70

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Broderick AC, Godley BJ, Hays GC (2001). Trophic status drives interannual variability in nesting numbers of marine turtles ProcRSocB 268:1481-1487

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Glen F, Broderick AC, Godley BJ, Metcalfe JD, Hays GC (2001). Dive angles for a green turtle (Chelonia mydas) JMarBiolAssUK 81:683-686

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Godley BJ, Broderick AC, Downie JR, Glen F, Houghton JD, Kirkwood I, Reece S, Hays GC (2001). Thermal conditions in nests of loggerhead turtles: further evidence suggesting female skewed sex ratios of hatchling production in the Mediterranean JExpMarBiolEcol 263:45-63

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Godley BJ, Broderick AC, Mrosovsky N (2001). Estimating hatchling sex ratios of loggerhead turtles in Cyprus from incubation durations MarEcolProgSer 210:195-201

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Hays GC, Ashworth JS, Barnsley MJ, Broderick AC, Emery DR, Godley BJ, Henwood A, Jones EL (2001). The importance of sand albedo for the thermal conditions on sea turtle nesting beaches Oikos 93:87-94

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Kasparek M, Godley BJ, Broderick AC (2001). Nesting of the Green Turtle, Chelonia mydas, in the Mediterranean: a review of status and conservation needs ZoolMidEast 24:45-74

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McGowan A, Broderick AC, Deemings J, Godley BJ, Hancock EG (2001). Dipteran infestation of loggerhead (Caretta caretta) and green (Chelonia mydas) sea turtle nests in northern Cyprus JNatHist 35:573-581

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McGowan A, Rowe LV, Broderick AC, Godley BJ (2001). Nest Factors Predisposing Loggerhead Sea Turtle (Caretta caretta) Clutches to Infestation by Dipteran Larvae on Northern Cyprus Copeia 2001:808-812

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Broderick AC, Godley BJ, Reece S, Downie JR (2000). Incubation periods and sex ratios of green turtles: highly female biased hatchling production in the eastern Mediterranean MarEcolProgSer 202:271-281

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Hays GC, Hochscheid S, Broderick AC, Godley BJ, Metcalfe JD (2000). Diving behaviour of green turtles: dive depth, dive duration and activity levels MarEcolProgSer 208:297-298

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Broderick AC, Godley BJ (1999). Effect of tagging marine turtles on nesting behaviour and reproductive success AnimBehav 58:587-591

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Godley BJ, Broderick AC, Moraghan S (1999). Short-term effectiveness of Passive Integrated Transponder (PIT) tags used in the study of Mediterranean marine turtles ChelonConservBiol 3:477-479

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Godley BJ, Thompson DR, Furness RW (1999). Do heavy metal concentrations pose a threat to marine turtles from the Mediterranean Sea? MarPollutBull 38:497-502

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Hochscheid S, Godley BJ, Broderick AC, Wilson RP (1999). Reptilian diving: highly variable dive patterns in the green turtle Chelonia mydas MarEcolProgSer 185:101-112

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Mckenzie C, Godley BJ, Furness RW, Wells DE (1999). Concentrations and patterns of organochlorine contaminants in marine turtles from Mediterranean and Atlantic waters MarEnvironRes 47:117-135

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Godley BJ, Gucu AC, Broderick AC, Furness RW, Solomon SE (1998). Interaction between marine turtles and artisanal fisheries in the eastern Mediterranean: a probable cause for concern? ZoolMidEast 16:49-64

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Godley BJ, Thompson DR, Waldron S, Furness RW (1998). The trophic status of marine turtles as determined by stable isotope analysis MarEcolProgSer 166:277-284

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Laurent L, Casale P, Bradai MN, Godley BJ, Gerosa G, Broderick AC, Schroth W, Schierwater B, Levy AM, Freggi D, Abd El-Mawla EM, Hadoud DA, Gomati HE, Domingo M,Hadjichristophorou M, Kornaraky L, Demirayak F, Gautier C (1998). Molecular resolution of marine turtle stock composition in fishery bycatch: A case study in the Mediterranean MolEcol 7:1529-1542

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Broderick AC, Hancock EG (1997). Insect Infestation of Mediterranean Marine Turtle Eggs HerpetolRev 28:190-191

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Godley, BJ, Smith, S. M, Clark, P. F & Taylor, J. D. (1997). Molluscan and crustacean items in the diet of the loggerhead turtle,Caretta caretta (Liiaeus, 1758) [Testudiies: Chelonidae] in the eastern Mediterranean JMollStud 63:474-476

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Broderick AC, Godley BJ (1997). Observations of reproductive behavior of male green turtles (Chelonia mydas) at a nesting beach in Cyprus ChelonConservBio 2:615-616

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Annette, C. Broderick & Brendan, J. Godley (1996). Population and nesting ecology of the Green Turtle, Chelonia mydas, and the Loggerhead Turtle, Caretta caretta, in northern Cyprus Zoology in the Middle East 13:27-46

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